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Stimulus Receptor Neural
Relay Cortex |
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In Vision |
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Light Rods/Cones LGN of
Thalamus Striate |
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In Audition |
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Sound Hair Cells MGN of
Thalamus Sup. Temp. G. |
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Other Generalities |
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Always more than one pathway in brain |
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Always more than one brain target |
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Ultimately sensory information is combined |
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Amplitude is the difference in air pressure
between the compression and rarefaction. |
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The measure of sound amplitude is the relative
measure called decibel or dB. |
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Where P = air pressure; P2 = power |
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dB SPL, P2=0.0002 dynes/cm2
which is near the absolute threshold for hearing. |
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Resonance |
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All physical mater will most easily vibrate at
certain frequencies. |
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This is true of our ear. |
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Thus some frequencies will more easily enter our
ear |
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It helps us determine the frequencies of
incoming sounds as we shall see. |
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The physical dimensions are related to but not
the same as the psychological dimensions: |
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frequency <> pitch |
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amplitude <> loudness |
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Three Major Divisions |
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Outer Ear receives sound directs it to the rest
of the ear. |
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Pinna - directs sound energy to middle ear and
helps perception of the direction. |
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External Auditory Meatus or Canal - 2.5 to 3 cm
long, 7 mm wide Resonates at about 2-4K Hz. |
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Tympanic Membrane |
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Middle Ear transmits sound information to inner
ear. |
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Ossicles transmit and amplify sound energy. |
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Malleus - Hammer |
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Incus - anvil |
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Stapes - stirrup |
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Eustachian Tube |
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Inner Ear is where transduction of sound
information occurs. |
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Cochlea (snail) with the |
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Oval Window |
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Round Window |
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The Cochlea - |
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Latin for seashell which is what it looks like |
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Basilar membrane runs most of the length of the
cochlea dividing in the top and bottom. |
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The base is right below the oval window where
the sound energy enters |
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The apex is at the other end. |
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Hair Cells are the receptors and run the length
of the Basilar Membrane in two sets |
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inner 1 row ~ 3500 |
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outer 3 rows ~20000 |
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Tectorial Membrane - across top of Hair Cells |
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Auditory Transduction |
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Transduction is the conversion of energy from
one form to another, e.g., sound pressure to neural impulses |
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The Traveling wave. |
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Wave set up by action of stapes on oval window |
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Point of Maximal Displacement depends upon the
frequency of the tone. |
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High Frequencies near the base. |
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Low frequencies near the apex. |
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The shearing force |
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The traveling wave bends the basilar membrane |
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This bends the hair cells. |
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The experience of sound most closely related to
amplitude or intensity. |
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Examples of sounds at different dB SPL levels
for comparison. |
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Rustling Leaves =~20 dB |
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Average Speaking Voice =~60 dB |
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Heavy Traffic =~80 dB |
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Rock Band =~120 dB |
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Pain/Damage Threshold =~130 to 140 dB |
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Loudness
differs in many ways from intensity. |
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The threshold depends upon intensity and
frequency. |
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Intensity doubles every 3 dB; loudness doubles
every 8 dB. |
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The dimension of sound that most closely relates
to frequency. |
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The higher the frequency the higher the pitch. |
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Discrimination between two pitches depends on
the frequency of the lower pitch: |
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Weber Fraction: (f1 - f2)/f2 =
0.004 |
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e.g. |
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(251-250)/250= 0.004 (1004-1000)/1000=0.004 |
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Pitch is not the same as frequency |
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Pitch will change as intensity is increase and
frequency is kept constant. |
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First studied by Fletcher and Munson (1933). |
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Called Fletcher-Munson Curves or Equal Loudness
Contours. |
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Method: |
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Subjects adjusted tone of different frequencies
to match loudness of 1000 Hz tone |
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the intensity of 1000 Hz tone was varied over
trials. |
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Thus, all tones that match a 1K Hz tone of a
given intensity should all be equally loud and connecting those on a graph
of intensity by frequency should give an equal loudness contour. |
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As intensity of the 1K Hz tone increase, the
contours get flatter. |
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Relates to the Loudness button on your stereo. |
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This relationship again illustrates the
difference between physical dimensions and psychological experience. |
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Sound Button on Stereo |
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Most recording are at region where loudness if
fairly constant across frequency. |
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We may play at a lot lower level where loudness
does depend on frequency |
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Alters what we hear because we lose sensitivity
to low and high frequencies faster than middle frequencies. |
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Sound button compensates for this by boosting
high and low frequencies. |
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A mathematical procedure to break down complex
waveforms in to simple components, usually sinewaves. |
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The ear does something like this. |
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Let us use this stimulus as our complex wave. |
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It is called a square wave. |
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The Frequency Domain |
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Frequency of Sinewave along the x-axis |
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Amplitude of Sinewave along the y-axis |
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Beats |
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Perception of intensity changes from two nearby
frequencies |
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From constructive and destructive interference |
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Frequency of beating is difference in frequency
between the two tones, e.g. 101-100 = 1 Hz beats |
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Missing fundamental |
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Fundamental is lowest pitch of a tone |
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higher frequencies called harmonics or partials |
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Perceive a same pitch even without fundamental |
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Allows us to tell female vs. male voices on the
telephone. |
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DEFINITION: one tone is rendered less
perceptible by another auditory stimulus. |
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Tone Masking |
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low tones will mask higher tones better. |
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due to shape of traveling wave (skewed towards
base, higher frequencies). |
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Noise Masking |
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Noise is sound energy that lacks coherence. |
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Beyond a point adding more frequencies to the
noise does not increase masking. |
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Critical bands: region of basilar membrane where
sound energy is summed together. |
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Consider Noisy Environments |
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How keep all the sounds distinguishable? |
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Consider sirens and other alerting sounds? |
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Is simply loud enough or necessary? |
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Eyes can see only in one direction at a
time. Ears are not so limited. |
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Interaural Time Difference/Phase |
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Description - sound has to travel farther to ear
on farther side of head |
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This difference can be detected if as small as
0.1 msec. |
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Works for clicks and tones with frequencies <
1000 Hz |
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Precedence Effect - Tendency to suppress later
arriving parts of a sound |
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Interaural Intensity Differences |
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Description - Head shadows sound so that farther
ear will hear a slightly less intense sound. |
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Just as we suppress later sounds, we suppress
less intense sounds. |
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Works best for relatively high frequencies. |
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This ability to
hear sounds from all directions is useful to design alerts. |
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The Detection Situation |
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The Stimulus is: |
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Subject |
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Judges |
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Stimulus |
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to be: |
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